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Nucleus                                        region in space where an electron is present most of the time.
                               (6 protons + 6 neutrons)                       The area in space where electrons circle around the nucleus is
                                                                              known as an  energy level   or    electron    shell.  The innermost

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                                   e -                                        energy level may contain one or two electrons. As you can see
                                               e -
                                                                              in    FIGURE 1.3 , hydrogen has one electron in the first energy


                                                                              level, which is also its only energy level. Many elements that
                              e -                              e -  Electron  are important in biology, such as carbon,  nitrogen, and oxygen,
                                                                              contain two energy levels. In these atoms, the second energy
                                       +                       +   Proton     level may hold up to eight  electrons.  Figure 1.3  shows how
                                    +
                                        +                                     carbon’s six electrons are placed in the energy levels: two are in
                                   +   +   +                       Neutron    the first energy level and four are in the second energy level.
                                                                                   The amount of energy in a level depends on its location.
                                                     e -                      Electrons closer to the nucleus have less energy and are less
                            e -                                               reactive than those further from the nucleus. Atoms are most
                                                                              stable when their energy levels are full. When electrons fill the
                                                                              energy levels closest to the nucleus, rather than ones further
                                  e -                                         away, the element becomes more stable. If an electron gains
                                                                              energy and moves or “jumps” to a level further away from the
                                                                              nucleus, the atom is less stable. As we will see later, molecules
                                   Carbon atom
                                                                              are often formed when atoms share electrons to fill their out-
                      FIGURE   1.2         A carbon atom                      ermost energy levels.

                      Most carbon atoms have six protons, which are positively charged;     Since chemicals often react to complete their outermost
                      six neutrons, which are neutral; and six electrons, which are   energy levels, simple diagrams like those in  Figure 1.3  give
                       negatively charged.                                    a sense of how many electrons an element must gain or lose
                                                                              to have a full outermost energy level. In the case of carbon, it
                            Typically, an atom has equal numbers of protons and   must gain four electrons in the outer shell for a total of eight
                      electrons. As you can see in  Figure 1.2  carbon possesses   electrons. Hydrogen may either gain one electron or lose an
                                                          ,
                      six positively charged protons and six negatively charged   electron to have its outermost shell complete. If an electron is
                                                                                                 +

                        electrons. The charges add up to zero, so this carbon atom is   lost, a hydrogen ion   (H )  forms because it now has one more

                      electrically neutral.                                   proton than electrons. We will study hydrogen in greater
                            Certain chemical processes cause an atom to either   detail when we discuss cellular energetics in Unit 3.
                      gain or lose electrons. An atom that has lost an electron is
                        positively charged, and one that has gained an electron is neg-     Chemical Properties of Elements
                      atively charged. Electrically charged atoms are called  ions.    The electrons in an element’s outermost energy level are

                      The charge of an ion is specified as a superscript to the right   known as the  valence electrons   .  The   periodic table of

                                                        +

                      of the chemical symbol. For example,    H   indicates a hydrogen
                      ion that has lost an electron and is positively charged. Posi-
                      tively charged ions are called cations and  negatively charged
                      atoms are anions.                                                                  In this simpli ed diagram, the
                                                                                                         electron energy levels are depicted
                         Electrons                                                                       as circles and the electrons that
                                                                                                         occupy them as dots. The cloud in
                        The movement of electrons from one molecule to another   Hydrogen                the center is the nucleus.
                      is the foundation of energy transfer in many biologi-                    Carbon
                      cal  reactions. Therefore, understanding electron transfer is         FIGURE   1.3      Energy levels for hydrogen and carbon



                      essential to the study of cellular biochemistry.        The hydrogen atom contains one energy level with a single electron,
                            Electrons move around the nucleus as a cloud of points   while the carbon atom has two energy levels with two electrons in the
                      that is denser where the electron is most likely to be. The   first energy level and four electrons in the second. The energy levels
                      exact path of an electron varies, but it is possible to identify a   are depicted as circles and the electrons as orange dots.


                                                                                                    MODULE 1   ELEMENTS OF LIFE   31

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